At any scale! Micro, meso, and macro scale.
"From the smallest, micro-scales to large, macro-scales, the workings of many systems can be understood at multiple levels. Growing evidence suggests that the mesoscale, which connects the two extremes, is a good starting point when trying to build an ontology or complete understanding of a system. ...
Most insect-cognition researchers agree on the macro- and microscale models that describe how insects navigate the world — the neurons and brain structures at the microscale and the behaviors and flightpaths at the macro. But ... the field has been divided for decades over what happens in between — how insects transpose environmental stimuli and memories into behavior. That divide, the authors write, “is best understood as a mesoscale disagreement.” ..."
From the abstract:
"Recent accounts of multiscale modeling investigate ontic and epistemic constraints imposed by relations between component models at varying relative scales (macro, meso, micro). These accounts often focus especially on the role of the meso, or intermediate, relative scale in a multiscale model. We aid this effort by highlighting a novel role for mesoscale models: functioning as a focal point, and explanation, for disagreement between researchers who otherwise share theoretical commitments. We present a case study in multiscale modeling of insect behavior to illustrate, arguing that the cognitive map debate in neuroethology research is best understood as a mesoscale disagreement."
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